What is the Parasympathetic System and its function?
Keywords: Craniosacral outflow| hypothalamus controls| limbic system| third |seventh|nineth| tenth
cranial nerve|
Edinger Westphal nucleus|Superior|inferior
salivary nucleus| pterygopalatine ganglion| esophageal plexus|gastric plexus|
The autonomic nervous
system is also known as the vegetative nervous system. It has two divisions: sympathetic
and parasympathetic.
1. The sympathetic system arises from all thoracic spinal
segments to the first two lumbar spinal segments, and sometimes from the third
lumbar spinal segment are also involved. Therefore this is also known as
thoracolumbar outflow.
2. The parasympathetic system arises from the brain and sacral
spinal segments, S2, S3, and S4. Therefore this is known as craniosacral
outflow.
Sympathetic and
parasympathetic systems usually act opposite to each other.
Table of Contents:
Sr. no |
contents |
1. |
Types
of Autonomic nervous system Sympathetic system Parasympathetic
system |
2. |
Parasympathetic
system |
3. |
Neurotransmitter Acetylcholine Adrenalin Noradrenaline |
4. |
The
hypothalamic controls The
limbic system control |
5. |
Effects
of the parasympathetic system on the
heart On
blood vessels On
metabolism On
digestive system Respiratory
system Genitourinary
system Nervous
system On
skin On
temperature regulation |
6. |
Fig
1 and2 |
The
hypothalamic control
The hypothalamuscontrols the sympathetic and parasympathetic systems.
The posterolateral
nucleus of the hypothalamus controls sympathetic activity and anteromedial
parasympathetic activity.
The posterolateral
nucleus stimulates sympathetic activity, and the anteromedial nucleus
stimulates parasympathetic activity.
Multiple areas of the
CNS influence the action of the anteromedial and posterolateral nucleus of the
hypothalamus via the limbic system.
The limbic system is
associated with the auditory cortex, visual cortex, parietal lobe, and
prefrontal lobe. In addition, the limbic system affects the activity of the
sympathetic and parasympathetic systems.
The parietal lobe is
concerned with memory, and the prefrontal cortex with the thinking process.
Therefore, higher centers will control the activity of the limbic system and
hypothalamic centers.
.
After interpretation of different pieces of
information coming to the limbic system, higher centers regulate the
hypothalamus and, therefore, sympathetic and parasympathetic activity.
The sympathetic
system is also known as the fight and flight or catabolic system, while the
parasympathetic system is known as the ‘rest and digest’ or ‘anabolic system.’
Cranial outflow
Parasympathetic nerves
arise from the brain stem –Midbrain, Pons, and Medulla -as well as from the
sacral spinal segments of the spinal cord; therefore, it is also known as
cranial sacral outflow.
The third cranial nerve
- Oculomotor
The seventh cranial nerve- Facial
The ninth cranial –Glossopharyngeal,
and
The tenth cranial nerve- Vagus,
They are the nerves through
which the cranial parasympathetic nerve emerges from the brain and is distributed
to the peripheral structures.
The third cranial nerve
The third cranial nerve is the
oculomotor nerve which contains somatic motor and parasympathetic nerves. The
somatic motor fibers arise from the third nerve nucleus present in the midbrain.
It supplies some of the extrinsic muscles of the eye. The parasympathetic motor
fibers arise from the Edinger Westphal nucleus of the midbrain and supply the
smooth muscles of the pupil and ciliary body of the eye.
The seventh cranial nerve
The somatic motor fibers of
the seventh supply the muscles of facial expression and the muscles of the
scalp. Its nucleus is situated in the pons. The sensory and parasympathetic fibers
of the facial nerve are together known as the nerve of Wrisberg. The parasympathetic
fibers from the superior salivary nucleus supply the submandibular and
sublingual salivary glands. The sensory fibers bring test sensation from the
anterior two-thirds of the tongue to the brain. The sensory fibers of somatic
sensation of the 7 th nerve ends at the ‘nucleus of tractus solitarius,’ and
from that, the final order neurons terminate on the superior salivary nucleus.
The ninth cranial nerve
The ninth cranial nerve is the
glossopharyngeal, which contains parasympathetic fibers arising from its
nucleus in the inferior salivary nucleus. Its somatic motor fibers are ‘special
visceral motor fibers’ arising from nucleus ambiguus ‘in the medulla, which
supply the stylopharyngeus muscle.
Sensory fibers of the
glossopharyngeal nerve end at
1. Those fibers with
bringing test sensation end at the ‘nucleus of tractus solitarius ‘
2. Those fibers bringing
general information ends at the spinal nucleus of the Trigeminal nerve, and
3. Those fibers bring baroreceptor
information from the carotid sinus end at the ‘dorsal motor nucleus of the vagus.’
The 10th cranial nerve
The 10th cranial nerve is
the vagus. It is the essential cranial nerve and contains the parasympathetic,
somatic motor, and sensory fibers. The parasympathetic motor fibers arise from
the dorsal motor nucleus of the vagus situated on the floor of the fourth
ventricle in the medulla. The fibers supply the thoracic viscera- heart and lungs,
and viscera of the upper
abdomen. The sensory fibers bringing information from these viscera end in
various parts of the brainstem. The somatic efferents arise from the nucleus
ambiguus and supply some laryngeal and pharyngeal muscles.
The parasympathetic
stimulation causes increased activity of gastrointestinal glands, stimulates
glands to increase secretion, and causes increased motility resulting in
increased movements of the GIT.
Parasympathetic nucleus
1. Edinger westphal
nucleus (3rd nerve ) is situated in the midbrain.
2. Superior
salivatory nucleus(7 th nerve) is situated in the lower part of the
pons
3. Inferior
salivatory nucleus (ninth nerve )is situated in the upper part of the medulla,
and
4. Dorsal nucleus of
the vagus (tenth nerve ) is situated in the medulla.
These nerves contain the parasympathetic
system fibers along with somatic fibers.
Outflow from the sacral segment
From the lateral horn
of the second, third, and fourth spinal segments of the sacral part of the
spinal cord, the parasympathetic fibers emerge to form ‘Pelvic splanchnic
nerves’ one on either side. It is
also known as ‘nervous erigens’(Nervous erigentis pl.).
Ganglions
They are preganglionic
fibers and relay in their ganglion, called the ciliary (3rd ), submaxillary(7th),
and otic(9th) ganglion. Ganglions for the vagus are situated close
to their target organs. The preganglionic parasympathetic fibers are myelinated
and long, while the postganglionic fibers are short and unmyelinated.
Nerve distribution of cranial
outflow
The parasympathetic nuclei
‘Edinger Westphal nucleus’ of the third cranial nerve supplies the intrinsic
muscles of the eye through the ciliary ganglion, an intramural ganglion. It
supplies the ciliary muscle and the sphincter pupillae.
1. When ciliary muscle
contracts, the suspensory ligaments relax, the lens becomes globular, and near
vision accommodation.
2. The other muscle is the
sphincter pupilli. When this contracts, the pupil constricts, the condition is
known as meiosis.
The parasympathetic nuclei
of the 7th cranial nerve are ‘Superior salivatory nucleus. The preganglionic
parasympathetic fibers ‘synapse
1. The pterygopalatine ganglion and its postganglionic fibers supply
lacrimal, nasal, and palatine glands. It causes increased secretion from these
glands.
2. In the submandibular ganglion, and the postganglionic fibers
supply the submandibular and sublingual salivary glands. It increases
secretions of the glands-watery, enzyme-rich saliva.
The parasympathetic nucleus
of the 9th cranial nerve is ‘Inferior salivatory nucleus. The preganglionic
fibers synapse in the otic ganglion, and the postganglionic fibers supply the
parotid gland. It increases secretions of the glands-watery, enzyme-rich
saliva.
The parasympathetic of the 10th
cranial nerve ‘Dorsal nucleus of the vagus’ supplies different body organs by
nerve plexuses. Their ganglions are very close to the organ.
The esophageal plexus supplies the esophagus and increases its peristalsis.
The gastric plexus supplies the stomach with anterior and posterior gastric nerves,
increasing its peristalsis, secretion, and absorption.
The small intestine, large intestine up to
two-thirds of the transverse colon, pancreas, liver, and gallbladder. The
parasympathetic of the 10th cranial nerve supply the smooth muscles and
exocrine glands.
The celiac plexus supplies the duodenum, jejunum, ileum, caecum, ascending colon,
and two-third of the transverse colon. The postganglionic parasympathetic
fibers increase motility, secretion, and absorption in these organs.
It increases urine
formation and motility of the ureter.
The cardiac plexus: The parasympathetic fibers are present in the
atrium and cause contraction of the atrium, in AV node delay the conduction. The
parasympathetic stimulation causes bradycardia and a fall of cardiac output and
blood pressure. Parasympathetic fibers are not present in the ventricle.
The pulmonary plexus: Parasympathetic
fibers in the lungs supply bronchial muscles, bronchial glands, and bronchial
arteries. The parasympathetic stimulation causes bronchial constriction, increases
mucus secretion, and reduces bronchial blood supply.
The hepatic plexus:
Parasympathetic fibers from the right vagus form the hepatic plexus. In the
liver, it increases glycogenesis, contraction of the gall bladder, and
relaxation of the sphincter of Oddi. In addition, it releases insulin from beta
cells of the ‘islet of Langerhans’ of the pancreas.
Cranial outflow |
The sacral spinal segments
S2,3,4 join to make ‘pelvic splanchnic nerve.
The postganglionic
parasympathetic fibers supply one-third of the transverse colon, descending
colon, sigmoid colon, and rectum and increase motility, secretion, and
absorption.
These fibers relax the
internal anal sphincter. (The external anal sphincter is under voluntary
control.)
The bladder contracts, and
the internal urethral sphincter relaxes. (The external urethral sphincter is under
somatic control).
It causes erection in males
and clitoris engorgement in females due to blood vessels dilations and
increased blood flow.
Sacral outflow |
We have seen that the organs
receive both sympathetic and parasympathetic nerves. Both are present, and the
resultant of the opposing effects determines the state of activity of a
particular organ at a specific time.
Hashtags: craniosacral outflow|Edinger Westphal nucleus|
esophageal plexus|gastric plexus|
Internal links: https://blog.totalphysiology.com/2021/08/hypothalamus-html
https://blog.totalphysiology.com/2021/08/hypothalamus-connections-and-functions.html
https://blog.totalphysiology.com/2021/11/what-is-autonomic-nervous-system-what.html
External link: http://en.m.wikipedia.org/wiki
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